Some of the winning images from the Wildlife Photographer of the Year competition.
Diana Rebman (US) photographed these gorilla twins in Rwanda’s Volcanoes National Park. She said: "The mum with twins was feeding and the silverback ran her off, he wanted the nettle she was eating. In my photo she looks stressed." She was only allowed one hour to take photos. "We were basically on a stopwatch, you have to take what you get," she added.
Chris Aydlett (US) spotted this raven perched on the south rim of the Grand Canyon. He thought the bird would fly away as soon as he got out of his car, so he grabbed his camera. But it stayed, watching him and giving him time to select a rock as a substitute tripod.
Andrew Walmsley (UK) was runner up in the mammal behaviour category. The critically endangered Celebes crested macaques are only found on Sulawesi and nearby islands in Indonesia. He was photographing the male who was gazing peacefully out to sea, when suddenly the peace was shattered by noise from behind. "I turned round to see these young males charging. They were screaming, kicking up gravel and making as big a show as possible, their faces full of expression."
At a waterhole in the Kalahari National Park, Peter Delaney (Ireland) came across scores of white-backed and lappet-faced vultures on an eland carcass, squabbling over the meat.
The overall winner of the Veolia Environment Wildlife Photographer of the Year is Greg du Toit (South Africa) for his photo Essence of Elephants. He said: "I’ve wanted to create an image that captures their special energy and the state of consciousness that I sense when I’m with them. This image comes closest to doing that."
Fourteen year-old Udayan Rao Pawar (India) is junior overall winner for a nesting colony of gharial crocodiles. "I could hear them making little grunting sounds," he said. "Very soon a large female surfaced near the shore, checking on her charges. Some of the hatchlings swam to her and climbed onto her head. Perhaps it made them feel safe." It turned out that she was the chief female of the group, looking after all the hatchlings.
This picture of an endangered Amur, or Siberian, tiger is one of only a very few taken in the wild without the use of a camera trap. Toshiji Fukuda (Japan) has been photographing wildlife in the Russian Far East for more than 20 years, so when he heard that tiger tracks had been found on the shore of Russia’s Lazovsky Nature Reserve, he knew this was his chance. His photo wins The Gerald Durrell Award for Endangered Species.
Isak Pretorius (South Africa) is winner of the Behaviour Birds category. He captured this lesser noddy caught in a colossal spider's net on the Seychelles island. He explained that it was "a bit of an eerie picture because the bird is still alive, you can see the pain and the inevitable story that’s evolving." Even if they struggle free, the silk clogs up their feathers so they can’t fly.