Seabirds nesting on the Farne Islands in Northumberland have been hit by heavy rain which killed many of their young, the National Trust has said.
Arctic terns, puffins, guillemots and shags all suffered losses as chicks including pufflings - baby puffins - were battered by nearly 5in (13cm) of rain in 24 hours on 13 June.
Gwen Potter, countryside manager for the National Trust, said: "The significant rainfall sadly caused many ground-nesting Arctic tern chicks to perish due to exposure to the elements.
"We don't know the full impact yet but estimate that tern numbers are likely to dip by up to 35% this year.
"We also know that 300 pufflings perished on one of the islands. Puffins are ground-nesting birds and unfortunately their burrows flooded. "