Presented by John Humphrys and Evan Davis.
Katherine Dyton, her daughter Carol and Lord Sutherland consider the provision of care for the elderly.
Security correspondent Frank Gardner reflects on his earlier interview with the Taliban's main spokesman, Zabihullah Mujahid, and discusses the relationship between Al Qaeda and the Taliban.
Dr Karen McComb explains how cats purr to manipulate their owners.
Hannah Clarke, a 16-year-old girl from Wales who made history as a baby when she had a donor heart grafted onto her own, has made a full recovery. Miss Clarke, her parents and Sir Magdi Yacoub, the pioneering surgeon who performed Hannah's original transplant, discuss the 'magic' recovery.
More than 20 Iraqis who worked for British forces are to sue the UK government, the BBC has learned. Angus Crawford reports.
Thought for the Day with Reverend Rosemary Lain-Priestley.
Chief Executive of the Independent Schools Council David Lyscom and Chairman of the Charity Commission Dame Suzi Leather discuss how the loss of charitable status affects tax benefits for independent schools.
Head of the Army General Sir Richard Dannatt reflects on the memorial held by troops in Helmand province for eight British soldiers killed in Afghanistan.
Science correspondent Tom Feilden and Professor Ian Jones analyse the threat posed by swine flu.
Jonathan Lennie, of Time Out magazine, and Roger Wright, director of BBC Proms and controller of Radio 3, discuss the prevalence of inappropriate applause after - and even during - performances.
One of classical music's leading conductors, Sir Edward Downes, has died along with his wife, Lady Joan Downes, after they travelled to Switzerland to take their own lives at the Dignitas clinic. Roger Wright discusses Sir Edward's life.
Health Secretary Andy Burnham and his shadow counterpart Andrew Lansley discuss whether asking older people in England to take out insurance to pay for long-term care is a fairer system than the current one.
Leading anti-apartheid campaigner Archbishop Desmond Tutu discusses his hope that Africa is beginning to turn its back on its repressive regimes and to see the prosperity that comes from accountable government.
Angus Stickler reports on allegations that MoD officials ignored safety alerts before three separate air crashes.
Former Lord Chancellor Lord Mackay and Terry Sanderson, president of the National Secular Society, discuss if Christians are being persecuted in the workplace and whether workers who are religious should be allowed to exercise their right to free speech.