Insight, analysis and expert debate as key policy makers are challenged on the latest news stories.
Presented by Evan Davis and Nick Robinson.
SNP deputy leader Nicola Sturgeon discusses the party's by-election win in Glasgow last night.
Secretary of State for Scotland Des Browne and Professor John Curtice of Strathclyde University offer their views.
Newspapers have railed against what they claim is a new privacy law in all but name after Max Mosley's victory against the News of the World. Rachel Atkins of the law firm Schillings discusses whether lawyers agree with the media.
The British Library Sound Archive has acquired the audio recordings of the Survey of Anglo-Welsh Dialects, a unique investigation into the local vocabulary, pronunciation and grammar of English in Wales. Robert Penhallurick of the University of Wales discusses the survey.
In Afghanistan, another British soldier has died and at least four have been injured in Helmand province. Alastair Leithead reports from the front line.
Thought for the Day with John Bell of the Iona Community.
A Qantas jumbo jet flying from London to Australia has made an emergency landing in the Philippines after a hole appeared in its fuselage. Passenger Olivia Lucas describes the pandemonium.
Barack Obama's speech in Berlin last night won cheers in Europe, but it was really designed to boost his electoral chances at home. Jonathan Beale reports from Washington on the reaction in the US to his tour.
Fraser Nelson of The Spectator and Kevin Maguire of the Daily Mirror discuss where last night's by-election defeat leaves Gordon Brown.