Hierarchical animals have what is known colloquially as a 'pecking order'. Animals at the top of the hierarchy aggressively dominate those lower down the ranks in order to retain privileged access to resources such as food, mates or a safe places to sleep. Males are often likely to dominate a pack, but there are some exceptions, as in spotted hyenas where females outrank males. In other cases there may be an alpha female in addition to an alpha male, as in wolves where she is the only one that will bear pups.
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Pecking order
In toque macaque society, highborn females can pull rank on subordinates for food.
In Sri Lanka, toque macaque society is run by a ruthless class system. Those born into high-ranking families have a massive head-start in life. If a highborn female wants something from a low-ranking female she can take it, literally taking food from her mouth. If the subordinate refuses to co-operate, she risks being beaten by the rest of the troop.
Sneaky mating
Low-ranking male toque macaques will adopt sneaky tactics in order to mate.
Toque macaque troops are ruled by a strict hierarchy in Sri Lanka. The rights to mate should only be held by the alpha male and he spends time recruiting allies to ensure this is the case. However, if a female sees potential in a low-ranking male as a future leader, it is a shrewd move to mate with him instead. Both will face severe punishment from the troop if caught attempting this but it may be the only chance for the low-ranking male to pass on his genes. So as not to raise suspicion, the pair will use very subtle glances, eyelid fluttering and teeth flashing in order to signal their interest to one another. When the alpha male and his allies are not looking, the pair will then sneak off for a quick secret liaison.
Dangerous liaison
Mating in secret is a highly risky business for a lone wolf.
Mating in secret is a highly risky business for a lone wolf.
Parallel pacing
Fallow deer bucks strut shoulder to shoulder to size each other up.
Fallow deer bucks strut shoulder to shoulder to size each other up.
Chillingham cattle
Semi-wild Northumberland cattle hark back to ancient breeds.
Semi-wild Northumberland cattle hark back to ancient breeds.
Hedgehog
Tasmanian devil
African wild dog
Antarctic fur seal
Badger
Brown bear
Coyote
Ethiopian wolf
Giant river otter
Grey wolf
Meerkat
Southern Elephant Seal
Spotted hyena
Stoat
Walrus
African buffalo
American bison
Argali sheep
Bighorn sheep
Blackbuck
Blue wildebeest
Cattle and aurochs
Dall sheep
Giraffe
Guanaco
Hippopotamus
Markhor
Milu
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Musk ox
Nubian ibex
Springbok
Walia ibex
Water buffalo
Rabbit
Brush-tailed rock wallaby
Eastern grey kangaroo
Aye-aye
Barbary macaque
Bonobo
Chimpanzee
Common woolly monkey
Crowned lemur
Eastern Gorilla
François' langur
Gelada baboon
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Grey mouse lemur
Hamadryas baboon
Human
Indri
Japanese macaque
Macaques
Madame Berthe's mouse lemur
Modern and early humans
Olive baboon
Phayre's leaf monkey
Pied tamarin
Ring-tailed lemur
Siamang
Silky sifaka
Tarsiers
Toque macaque
Tufted capuchin
Verreaux's sifaka
Western gorilla
African bush elephant
Asian elephant
Columbian mammoth
Forest elephant
Woolly mammoth
Arctic ground squirrel
Bank vole
Black-tailed prairie dog
Brown rat
Capybara
Damaraland mole rat
European beaver
Grey squirrel
Gunnison's prairie dog
Naked mole rat
Patagonian mara
Common bottlenose dolphin
Indo-Pacific bottlenose dolphin
Sperm whaleA dominance hierarchy (in humans: social hierarchy) is the organization of individuals in a group that occurs when competition for resources leads to aggression. Schjelderup-Ebbe, who studied the often-cited example of the pecking order in chickens, found that such social structures lead to more stable flocks with reduced aggression among individuals.
Dominance hierarchies can be despotic or linear. In a despotic hierarchy, only one individual is dominant, while the others are all equally submissive. In a linear hierarchy, for example, in the above cited pecking order of chickens, each individual dominates all individuals below him and not those above him.
Dominance hierarchies occur in most social animal species that normally live in groups, including primates. Dominance hierarchies have been extensively studied in fish, birds, and mammals. Dominance hierarchies can be simple linear structures, which often arise from the physical differences among individuals in a group in relation to their access to resources. They are also influenced by the complex social interactions among individuals in the group.
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