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Part 1
Boyd receives an anonymous package - a Nazi ceremonial dagger. Attached is a number that refers to a cold case file - the unsolved murder of a young girl found stabbed to death on an East London street in May 1945.
Boyd discovers that the last person to look in the file was his old colleague Mel Silver, two weeks before her death. The pendant sent to Mel was evidence from the case.
The team learn that the dagger was bought from a dealer in war memorabilia by a woman called Sarah, the same woman who was in touch with Mel. They go to a house to talk to the Polish builder who took the dagger to the dealer to be sold. He's not there but Spencer questions his brother, Viktor. He shows Boyd and Spencer where he and his brother Jakub found the dagger, hidden behind a wall.
Outside Boyd confronts Sarah, who has been following him. She tells him she is working for Interpol on war crime cases and that the cold case relates to a case she is investigating about World War II crimes. She asks to be allowed to hook up with the cold case team but Boyd refuses.
Simon Dusniak, owner of the house, is shocked to discover that his mother, Abigail, has returned to the UK from her home in Panama. She challenges him as to why he has put her father Marek in a nursing home.
The team go back to the dealer's shop but find it ransacked and the dealer dead. It seems that Jakub came back to get the dagger and in the struggle accidentally killed the dealer.
Boyd finally lets Sarah tag along on the investigation. She and Spencer follow the Dusniak's family solicitor Dennis Holland to a retirement home. They then see Simon come out, and they follow him. They see that he is being watched by Jakub Cyrak. They go to arrest Jakub but he escapes.
In the retirement home Abigail gets Holland to witness Marek signing over all powers of attorney to the family money - even though the elderly Marek is barely aware of what he is doing.
When questioned, David and Simon lie about Marek, saying he's dead, in order to avoid troubling the old man. But Sarah and Spencer know that Marek is alive - and living in the retirement home.
Boyd gets Simon and David back in for questioning and then goes with them to the retirement home to question Marek. But when the team confront Marek with the SS dagger, Marek is thrown into confusion and asks for his medication. He takes a pill and as his body arches in pain, Boyd realises he's taken a cyanide pill to kill himself.
Back at the lab, Eve finds a military tattoo on the dead Marek's torso - this number identifies him as Nazi doctor Martin Heiss, who worked for Josef Mengele in extermination camps in Poland during the second world war, exterminating non-Aryan children.
Part 2
Dennis Holland admits to Boyd that Abigail forced him to witness Marek signing over powers of attorney to her.
Sarah and Grace talk to the Dusniak family rabbi about the family's history. They find the rabbi instructing Simon's son Ben before his forthcoming bar-mitzvah. Ben leaves but then Simon comes into the synagogue saying he can't find Ben.
Eve and Spence search the house at Chesterhill rd and find five bodies buried under the floor. Four of the bodies have a DNA match to the dead girl from the cold case (Bernadetta Dusniak) but one of the female children's bodies doesn't.
Grace questions David Dusniak and forces him to access long-repressed memories. David recalls with horror how he helped to bury the bodies in the house, and that his father is Josef Mengele.
Eve finds the same military numbers she found on Marek, on the unidentified little girl's corpse from the house.
Boyd and Sarah question Viktor Cyrak and he leads them to his brother Jakub. Simon comes into CCHQ to say that his son Ben is still missing and that he's seriously worried. Boyd suspects that Jakub's abducted Ben but he tells them that he's arranged to meet Abigail later that day and that she's going to pay him for the other documents he found behind the wall in the house.
When Jakub and Abigail meet, Boyd and Sarah are there and they arrest Abigail, who they now feel certain has taken Ben.
Abigail initially denies having Ben. The interview is interrupted by Eve and Sarah - they have new information that David was a 'lebensborn' child - part of a programme run by Mengele to perpetuate the Nazi \ Aryan bloodline.
Boyd goes back into the interview and accuses Abigail of killing the third child, Gabby, who was not, as she thought, another 'lebensborn' child, but in fact her real sister. Abigail breaks down and tells them where they can find Ben - in the boot of a car in a London car park. She had intended to smuggle him back to Panama to carry on the Nazi bloodline.
Ben is reunited with his father Simon.
Boyd is in New York with Sarah. They kiss.
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